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Here's my day so far: went to jail, lost the girl of my dreams and got my butt kicked pretty good. Still, things could be a lot worse. Oh, that's right... I'm falling to my death. Guess they can't. How did it all come to this? Well, my end starts at the beginning... The very beginning!
~ Megamind's opening monologue.
Oh, I'm shaking in my custom baby seal leather boots!
YOU DARE CHALLENGE MEGAMIND?!?!
~ Megamind, finally confronting Tighten as a hero.
Megamind is the titular main protagonist of the Megamind franchise, serving as the titular main protagonist of the DreamWork's 21st full-length animated feature film of the same name and the titular main protagonist of its short video.
He starts out as a villain and has always lost to his archenemy Metro Man, but when Metro Man fakes his death, Megamind creates a new hero to battle in his place named Tighten, but Tighten turns out to be evil and kidnaps and tries to kill reporter Roxanne Ritchi (who Megamind had feelings for). Megamind, with the help of his sidekick Minion and the Brainbots, saves Roxanne and Metro City from Tighten and becomes a hero. A disguised Metro Man congratulates Megamind for what he has done.
At first, Megamind enjoyed being a supervillain, as this was what was expected of him from most of society. He's grown this way due to being rejected and tormented by his peers throughout most of his childhood, and often got in trouble whenever he tried to fend for himself. As a result, he started to believe that being bad was all he was really good at in life, and became a supervillain.
Despite his intelligence, Megamind is bumbling, egotistical, clumsy, and while making his life dependent on his villain status, had somewhat of a naive idea of what it truly means to be a villain. For example, he plays up his status as a villain as more of a game rather than as an actual battle between good and evil, and believed that actual villainy simply amounts to simple crimes like robberies or vandalizing public property. This his further shown after believing he has finally killed Metro Man, where he realized that without a hero to fight, he also had no purpose in life, and therefore created Tighten in order to have someone even out his villainy.
Even as a self-proclaimed supervillain, he still showed a strong aura of morality within himself. Despite all the damage he purposefully caused to Metro City on a daily basis, his intention was never to harm any of the civilians, not even Roxanne Ritchi, who he'd kidnap regularly, or Metro Man, who he only thought to have killed accidentally. Ironically, it has even gotten to the point where the civilians aren't genuinely afraid of him, but instead rejecting him just for what they perceive him to be. He is even disgusted when Tighten refuses to use his powers for good, instead using them for his own personal gain, and requesting Megamind to join in on his evil plans.
After seeing all of the damage Tighten is causing to Metrocity, as well as his plan to kill Roxanne, Megamind suddenly has a change of heart after Metro Man suggests to him to take his place as the city's new hero, and when he finally gives a sincere apology to the Warden, who is actually Minion in disguise.
The DVD commentary of the film note that his costume and showmanship are purposely evocative of Alice Cooper.
Ben Stiller (who previously voiced Alex in the Madagascar franchise) and Robert Downey Jr. were both approached for the role of Megamind, but it turned down due to scheduling conflicts. However, Stiller appears in the role as the curator named Bernard.